Researcher Armor (unlock: 20 Research Computers standing at once): +15% wave research, +10 team research per 100 squad kills, +200 research at spawn.
Turrets
Laser Turrets can convert (250 research, pick one):
Gatling Turret: 40 damage at 10 shots/s, twice the power draw.
Heat Ray Turret: sweeps an 8 m line every 2.5 s for 1000 damage a second.
Gatling upgrades (100 each): Targeting Computer +40% range. Overdrive Barrels spin up to +50% fire rate during unbroken fire.
Heat Ray upgrades (100 each): Hotter Beam +250 damage a second. Armor Melt ignores armor. Goo Burner burns the Goo when no Buggos are in range.
Laser Turrets no longer have levels.
Buildings and Modules
New module: Critical Hit (Common). +5% chance to deal 5x damage. Copies stack.
The + button adds a module slot to a building (100, 200, 300, 400 research; up to 5). Levels no longer add slots.
Walls get 3 upgrades: +200 health (100 research), +300 health (200), +15 armor (200).
New Robotics Bay upgrade: Defibrillators (100 research). Robots run to downed players anywhere on the map and revive them at 50% health. The run goes on while medic robots are out.
Drop Shop: buy a module credit for 200 research.
Squad
Squad pings: middle mouse marks a spot for your squad.
While down, left click switches the camera between teammates and medic robots.
Buggos
New boss: Boom Slug. Lobs Boom Slugs at your base, then charges the Command Tower and explodes for 500.
Wasp boss: new moves. Energy Drain, Stinger Rain, Crash Dive and Hive Call.
Buggos attack any building that blocks their path.
Buggos no longer freeze trying to reach buildings they can't get to.
Other
Weapon tooltips show damage, attack speed, range, magazine and reload.
Tutorial: new steps for opening your bag and socketing a module.
Right-click cancels a building placement in one press.